http://elfquest.com/comic_viewer.php?fd=/gallery/OnlineComics/OQ/OQ20/_Original%20ElfQuest%20-%2020_page=1#_26# Wendy made Timmain word it this way: "small, ground-digging apes" and "winged insects". We have only the familiar vocabulary, not a biolocical linage. As long as there isn't another or more detailed explanation I take the literal meaning: Trolls developed from a homeworld creature that was close enough to our apes to call them so. And Preservers come from insects. Rodents were not mentioned. What are you up to, Potoo? Might be easier to talk when we know more.